When and how does environment impact the evolution of galaxies? We will approach this question by considering two extreme environments. First: The cores of massive clusters. Here, the largest, reddest galaxies of the local universe are found, Brightest Cluster Galaxies (BCGs). These galaxies are found mixed with diffuse intracluster light (ICL) and often on top of the cooling intracluster medium (ICM). We'll explore recent results from an integral field unit survey of local cluster cores which provides photometric and spectroscopic evidence of a break in age, between the old red and dead BCG cores, and the ICL that surrounds them. Second: Cluster scale filaments. Here, galaxies find themselves in moderate density environments, where they are potentially able to interact with each other and the intracluster medium, initiating starbursts and undergoing quenching. Recent SITELLE observations from CFHT provide clear examples of blue starforming galaxies in cluster feeding filaments. We'll discover how within these environments, the galaxy’s ability to interact with the surrounding medium and surrounding galaxies has the potential to drive its own evolution.